Shoulder Hump:
The hump is where a mass of muscles attach to the bear's backbone and give the bear additional strength and power.
P.Atrox : Absent
U.Arctos : Present
Edge : U.Arctos
Scapula Shape Index:
It's scapula width/forelimb length ratio.Higher score indicates greater muscle attachement on the shoulder joint as well as greater ability to make certain movements like full-functional upper extremity movement including protraction,retraction,elevation,depression,upward rotation,and downward rotation all else being equal.
P.Atrox : 21%
U.Arctos : 26%
Edge : U.Arctos
Scapula Length Index:
Scapula length divided by forelimb length. Higher score Indicates greater speed of movement via relative proportions of components of the proximal forelimb and exertion of greater force.
P.Atrox : 30%
U.Arctos : 30%
Edge : Draw
API:
Acromion process length/Forelimb length.Higher score in this ratio increases attachment for the deltoid muscle.
P.Atrox : 2%
U.Arctos : 6%
Edge : U.Arctos
SMI:
Deltopectoral crest length divided by humerus length. Higher score Displays greater mechanical advantage of deltoid and pectoral muscles at the shoulder joint.
P.Atrox : 62%
U.Arctos : 72%
Edge : U.Arctos
HRI:
It is humerus mediolateral diameter at mid-shaft/humerus length.Robust forelimb bones protect the animal from violent stresses during grappling.
P.Atrox : 10%
U.Arctos : 11%
Edge : U.Arctos
HRI2:
It is humerus anteroposterior diameter at mid-shaft/humerus length. Robust forelimb bones protect the animal from violent stresses during grappling.
P.Atrox : 16%
U.Arctos : 12%
Edge : P.Atrox
HKX:
A better score signifies relatively greater cortical thickness of the humerus.
P.Atrox : exact value unknown
U.Arctos : exact value unknown
Edge : U.Arctos
HKY:
A better score signifies relatively greater cortical thickness of the humerus.
P.Atrox : exact value unknown
U.Arctos : exact value unknown
Edge : U.Arctos
HCMI:
It is Humeral circumference/articular length Often-used variable in analyses of bone strengths in locomotion.The higher the ratio is, the thicker the Humeral bone is.
P.Atrox : 29%
U.Arctos : 34%
Edge : U.Arctos
HEI:
The HEI index measures the relative size of wrist stabilizing muscles and well as several grasping muscles in area which facilitate prey grasping. These muscles also likely help stability, pushing, and agility when legs are on ground.
P.Atrox : 27%
U.Arctos : 30%
Edge : U.Arctos
HTI:
It's Humeral trochlear length/Humeral length. Higher score increases muscle attachement on
the elbow joint and allows for more ulnohumeral flexion and extension.
P.Atrox : 20%
U.Arctos : 21%
Edge : U.Arctos
Brachial Index:
This measures radius length/humerus length. Lower scores in this ratio indicate greater
forelimb muscle mechanical advantage and results in stronger muscles all else being equal.
P.Atrox : 94%
U.Arctos : 86%
Edge : U.Arctos
Brachial Index 2:
This measures ulna length/humerus length. Lower scores in this ratio indicate greater forelimb
muscle mechanical advantage and results in stronger muscles all else being equal.
P.Atrox : 113%
U.Arctos : 96%
Edge : U.Arctos
Forelimb Proportion Index:
Length of proximal forelimb relative to length of distal forelimb ((Scapula length + Humerus length)/(Radius length + Metacarpal length)). Higher score Indicates greater degree of
morphological specialization for producing large out-forces in the forelimb.
P.Atrox : 132%
U.Arctos : 154%
Edge : U.Arctos
RRI:
This measures radius mediolateral diameter at midshaft divided by radius length. As mentioned above for humerus robusticity, a robust radius resists stresses on bones during fights and increases resistance to bites to the forelimb.
P.Atrox : 11%
U.Arctos : 11%
Edge : Draw
RRI2:
This measures radius anteroposterior diameter at midshaft divided by radius length. As mentioned above for humerus robusticity, a robust radius resists stresses on bones during fights and increases resistance to bites to the forelimb.
P.Atrox : 7%
U.Arctos : 6%
Edge : P.Atrox
OLI:
Olecranon process length divided by functional length of the ulna. Higher score Indicates greater relative mechanical advantage of the triceps brachii and dorsoepitrochlearis muscles used in elbow extension.
P.Atrox : 16%
U.Arctos : 19%
Edge : U.Arctos
URI:
It is ulna mediolateral diameter at mid-shaft/humerus length. As mentioned above for humerus
and radius robusticity, Robust forelimb bones protect the animal from violent stresses during
grappling.
P.Atrox : 6%
U.Arctos : 6%
Edge : Draw
URI2:
This measures ulna anteroposterior diameter at midshaft divided by ulna length. As mentioned above for humerus and radius robusticity, a robust ulna resists stresses on bones during fights and increases resistance to bites to the forelimb.
P.Atrox : 10%
U.Arctos : 10%
Edge : Draw
MANUS:
Manus digit 3 proximal phalanx length divided by metacarpal 3 length.Higher score Indicates greater relative proportions of proximal and distal elements of the manus and size of the palmar surface.
P.Atrox : 49%
U.Arctos: 55%
Edge : U.Arctos
Foot Posture:
All else being equal, species that are able to adopt plantigrade foot posture, such as bears,badgers, wolverines, many rodents, spider monkeys, and great apes, should be able to apply substantially greater free moments to the ground than species that are restricted to digitigrade or unguligrade foot posture. Relative to digitigrade foot posture, plantigrade posture improves performance in lateral striking, lateral pushing, downward striking,forward pushing and rearward pulling.
P.Atrox : Digitigrade
U.Arctos : Plantigrade
Edge : U.Arctos
CLAW:
Manus digit three terminal phalanx length divided by pes digit three terminal phalanx. Higher score is to be expected in species whose claws are more capable of breaking soil.
P.Atrox : 98%
U.Arctos : 136%
Edge : U.Arctos
Studies Used For The Above Comparison:
https://www.jstor.org/stable/23735739
As can be seen, most ratios point out to the Brown Bear having proportionally stronger shoulders, thicker Humeri,larger epicondyles,trochleas,olecrana, shorter forelimbs, longer proximal forelimbs, greater Manus Proportions, better Foot Posture and deadlier claws while the American Lion simply had more robust radii. Overall the Brown Bear was a stronger,more robust and better grappler than a similarly sized Atrox.Therefore, the Latter needs to significantly outweigh the Ursid to gain a considerable chance which would not be the case if we talked about Kodiak Bears and Coastal Grizzlies as some weight data suggest average weights of 352 and 334 kg respectively :
https://www.jstor.org/stable/3872615
The American Lion on the other hand merely weighed around 260 kg :
Taking the weight difference into account, it appears that Panthera Atrox wouldn't do much against a Coastal Grizzly or Kodiak Bear in a fight !





